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These join Argonaute-1 to regulate gene expression 89. Piwi-interacting RNAs are restricted to the gonad, and at least a subset of these arises by Piwi-catalysed cleavage of single-stranded RNAs 12 Here we show that Drosophila generates a third small RNA class, endogenous small interfering RNAs, in both gonadal and somatic tissues. Production of these RNAs requires Dicer-2, but a subset depends preferentially on Loquacious 145 rather than the canonical Dicer-2 partner, R2D2 ref. Endogenous small interfering RNAs arise both from convergent transcription units and from structured genomic loci in a tissue-specific fashion.

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Forstemann, K. PLoS Whitee.

Lee, Y. Nature— Denli, A. Processing of primary microRNAs by the Microprocessor complex. Saito, K. Jiang, F. Genes Dev. Howorth, St.

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